Here's a very interesting graph from the eurogenes blog:
https://drive.google.com/file/d/0B9o3EY ... view?pli=1
Basically, these are genetic profiles of various individuals.
I think that what it shows are two main gradients. The first gradient goes from the black-cross Neolithic samples in the centre-left to the pink-inverted-triangle Corded Ware sample above-right-of-centre. This is Europe. It represents a gradient between the Neolithic farmers at bottom left and the IE invaders at top right, and is largely geographic. From Corded Ware leftward we get east slavic, baltic west slavic/south slavic/germanic, and then romance. The cluster of pink romance samples right next to the Neolithics are all (as seen on a more detailed version elsewhere) Sardinians, who pretty much represent the original Neolithic population.
[The blue dot on the far left is Oetzi, from the chalcolithic. Apparently it's a general thing that the late neolithic and chalcolithic samples tend to be even less IE than the earlier neolithic ones. This may be selection out of some shared genetic elements, or may have something to do with suggestions of an overturning of the population in the mid to late neolithic, which didn't greatly affect the material culture - perhaps some specific (and hence out-of-the-average) population of europe-wide rulers or traders that emerged from a neolithic society? Or a resurgence of a surviving mesolithic line that skewed the averages? Anyway, the scale is clearly much smaller than the later IE invasion!]
The second gradient goes from the bottom left - who are the Bedouin - up through various egyptians, jews and palestinians, then up into the iranians and the caucasians. You can see the Armenians and the Persians in red and blue in that line, and the grey stars at the far end of it are East Iranians. This is the gradient between pre-IE middle easterners and the IE invaders, although it's also possible that the gradient was there before them - for instance, it may ultimately be a gradient between Afroasiatic invaders (represented by the Bedouin) and the native ME population, represented by the Caucasians. More on this later...
The bottom gradient then points very neatly at a green dot at the far right. These are the Yamnaya people, who are assumed to be the closest we have to the Proto-Indo-Europeans. So the gradient fits nicely with the idea of an IE invasion from the Pontic steppe into the middle east.
The top gradient doesn't exactly point at the Yamnaya. However! See the little do way away at the top left? Those are Western Hunter Gatherers - the mesolithic population of Europe. Imagine a line from the Neolithics to the Yamnaya, and then imagine that line being PULLED by the influence of those WHGs. That neatly drags it out of direct alignment with the Yamnaya.
This leaves only two things to account for. The big blob in the middle? That's the later mediterranean interchange, combining the two gradients. The people in that blob are greeks, western turks, jews, cypriots, maltese and sicilians. Notice also how, eg, south slavs are pulled in that direction too, and the romance dots nearest the bridge are italians.
The other thing is, what's that little nobble of grey dots at the top of the european gradient, pointing up toward the Mesolithics? Why, that's the Basques! Mostly representative of the same Neolithic/Yamnaya gradient as everybody else, but distinctly pulled toward the Mesolithic population. As you'd expect from the atlantic seaboard, if the neolithic farmers didn't quite reach them!
It's really all too perfect!
[Oh, and the other grey dots in Europe? The Finns, Estonians, and Russian Uralics are the ones near to the Slavs. The ones closer to the Germans are the Hungarians. Clearly the Uralics are part of the same gradients as the IE - whether that means a shared origin, or just influence from earlier inhabitants of these regions, I don't know]
Now, here's an even neater graph. It's off the same thing, as you can see, but a bit twisted around and some different samples. Same guy over at Eurogenes:
https://drive.google.com/file/d/0B9o3EY ... view?pli=1
What we see here are in fact three very clear gradients. Four if you include the Mediterranean interchange.
The big addition to this graph is the bright red dot - that's a neolithic Turkish woman. See how there's a nice gradient between Neolithic Turkey and the 'WHG' (western hunter-gatherer - pre-invasion Europeans)? That gradient has the Early Neolithic Europeans at the bottom, then the Middle Neolithic Europeans at the top, as they incorporate more and more genes from the native population. Note how the Sardinians are in the middle of that gradient - isolated on Sardinia, they absorbed relatively less of the WHG genes, and have been mostly isolated ever since. They therefore probably represent more or less the "original" Cardial culture. [Apparently they're genetically something like 80-85% Neolithic Farmers]
Then of course you have the gradient from the Neolithic farmers (minus a gap, because they stopped being an influence and everyone else has drifted away from them over time) through the modern europeans, to NE europe, Corded Ware, modern Volga/Urals, and finally Yamnaya.
Then below that you've got the gradient from the bedouin, through the arabs, turks, iranians, armenians, caucasians, up to the tajiks, and then turning down toward the pashtuns. This doesn't point as neatly to the Yamnaya, although you could see the Tajiks as a Yamnaya + Caucasus mix.
Also of interest, though: the Yamnaya are about equidistant between the Caucasians and a little dot to the right called the EHG - the eastern hunter gatherers, the people in eastern europe and the steppe before the neolithic. Now, that could be coincidence. But apparently the analyses suggest that the best explanation of the Yamnaya is a cross between EHG and a group genetically close to Kartvelians. Which is nice, if you think that PIE looks sort of Caucasian...
[for the sake of completeness? Follow the line from the Tajiks through to the Pashtuns and then on in that direction - the IE population of India is somewhere off down there.]
[Oh, and note how ALL the WHG, SHG and EHG - all the mesolithics - are way off miles away from everyone else, but themselves in a nice enough little gradient]
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SO WHAT DOES THIS MEAN!?!?!?!?
Well, a nice model, from what i can see (and I'm no geneticist!) would be that:
- there was a gradient of quite different populations in the mesolithic, as we might expect.
- a near eastern population invaded and took over europe almost entirely, although the Basque were the least affected.
- these invaders were more closely related to modern arabs and jews, but not VERY related to them at all - later Afroasiatic invasians have clouded them over. In fact, you coudl squint and say that the modern middle east sort of looks like a three-way merger between Semitic invaders, Caucasians, and the 'original' neolithic farmers. That might fit neatly with history.
- meanwhile, caucasians also moved north into eastern mesolithic territory. Their ancestors were the Yamnaya
- the Yamnaya then invade Europe. Genetically their influence is much more felt in the east, of course, but ALL europeans, other than the Sardinians, are affected to some degree.
- meanwhile, some yamnaya also move south into the Iranian/Caucasian sphere. There's no surviving intermediaries here which might be just because of later slavic and iranian conquests. But given both the gap and the angle of the gradient, it may be that the Iranians were more of a ruling warrior class than the tribes who moved into europe.
- later, a branch of those Iranians/Bactrians/etc, part Yamnaya and part native, then pushed into India, again mostly as a warrior class. India at the time had entirely different genetics from either europe or the middle east.
- later, populations in southeast europe and the med gradually intermingle with populations from north africa and the middle east
And the relevence of this to the topic? No idea... Seemed like a good idea when I started?
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Oh, except that genetically it rather looks like there is no chance whatsoever that the Old European Hydronomy did all of these things:
a) exist
b) be particularly related to IE
c) be the LBK language.
You can pick any two of the three: IE-like LBK language that didn't exist (just overfitting by one philologist), real IE or IE-like language not related to LBK (an early wave of IE - perhaps a 'Corded Ware' language? or a 'Funnel Beaker' language?), or real LBK language not related to IE. That's because the presumed IE speakers - Corded Ware and Yamnaya - are bloody miles away genetically from any Neolithic european genes we've found at all.
Of course, it IS possible. Language could spread faster than genes, so an LBK-like language could have spread to the Yamnaya, either directly or from a Caucasian ancestor. But that seems much less probable than the alternatives!